DNA tests have confirmed that a Bulgarian couple are the biological parents of Maria, the girl found in a Greek Roma camp.
The case of Maria has drawn global attention, playing on the possibility of children being stolen from their parents or sold by them.
Charges have been brought against the couple who Maria was found living in the Roma settlement and they are being held in custody pending trial. They are also suspected of fraudulently obtaining birth certificates for a total 14 children.
The couple insist they were looking after Maria with their own five children after an informally-arranged adoption. The girl was placed into the care of a children's charity and her DNA details were provided to Interpol.
The woman who has now been confirmed as a DNA match for Maria was served with preliminary charges of child selling, but has not been detained.
While separately, police have arrested a childless couple in Greece on suspicion of buying a baby Roma girl and trying to register her as their own.
The couple were arrested in Athens after they allegedly paid a Roma woman €4,000 for the baby, a Greek police statement said. Authorities are looking for the baby's birth parents.
The suspects, aged 53 and 48, were expected to be charged with child abduction. Under Greek law this includes cases where a minor is voluntarily given away by its parents outside the legal adoption process.
And on Wednesday, another Roma couple was charged with child abduction on the eastern Greek island of Lesvos. Police found them with a baby boy that was not their own.
The couple allegedly told authorities that they were childless and had been given the baby by a Roma woman in Athens who had five children.