Grumpy Cat - one of the Internet's most recognisable feline stars - has died aged seven.
Grumpy, also known as Tardar Sauce, went viral in 2012 after photos of her famous frown spread online.
Her iconic expression quickly turned into a meme.
At seven years of age, the cat from Arizona was worth an estimated $1 million dollars.
Some days are grumpier than others... pic.twitter.com/ws209VWl97
— Grumpy Cat (@RealGrumpyCat) May 17, 2019
In a statement, Grumpy Cat's owners said: "Despite care from top professionals, as well as from her loving family, Grumpy encountered complications from a recent urinary tract infection that unfortunately became too tough for her to overcome.
"She passed away peacefully on the morning of Tuesday, May 14, at home in the arms of her mommy, Tabatha."
They add: "Her spirit will continue to live on through her fans everywhere."
Grumpy Cat's fame began with a Reddit post, with owner Tabatha Bundesen's brother Bryan posting an image of the kitten.
In 2013, the cat - then only 11 months old - made her first official public appearance at the SXSW festival in Austin, Texas.
Bryan Bundesen told CNN the cat's fame was 'accidental', explaining: "We thought it would get some laughs, with the Internet liking cats. And it just kept going."
The cat's Internet fame even led to a TV film called Grumpy Cat's Worst Christmas Ever - with the feline voices by Parks & Recreations star Aubrey Plaza.